Enzyme

Isolated from the plant Gentiana triflora. Transfers the hydroxycinnamoyl group only to the C-5 glucoside of anthocyanin. Caffeoyl-CoA, but not malonyl-CoA, can substitute as an acyl donor.

5AT

Enzyme

Anthocyanin 5-aromatic acyltransferase

Enzyme name

EC:2.3.1.153

Entry

Transferases;Acyltransferases;Transferring groups other than aminoacyl groups

Class

4-hydroxycinnamoyl-CoA:anthocyanidin 3,5-di-O-beta-D-glucoside 5-O-glucoside-6'''-O-4-hydroxycinnamoyltransferase

Sysname

4-hydroxycinnamoyl-CoA + an anthocyanidin 3,5-di-O-beta-D-glucoside = CoA + anthocyanidin 3-O-beta-D-glucoside 5-O-beta-D-(6-O-4-hydroxycinnamoylglucoside) [RN:R05163]

Reaction

CoA [CPD:C00010];anthocyanidin 3-O-beta-D-glucoside 5-O-beta-D-(6-O-4-hydroxycinnamoylglucoside) [CPD:C06363]

Product

Species information

Euphorbia lathyris

Species name

Euphorbia lathyris is a species of eudicot in the family Euphorbiaceae (spurge family).

Species info

Gene sequence info

Euphorbia_lathyris.5AT.fa

Download
>Casp05102.t1
MLEVCRVTPFTNSSTPATELTLPLRYFDMMWHSLSLTLTHFLPVAGNIIWPVDTAKPHVLYTPNDSVSFIIAESDSDLTHLSRNEIQEAADLHHYIPELHVSDTKAAIIAFQITFFPSQGFSIGISTHHGVFDGKSVTIFMKSWAHICKENFDDDVAALLPRDLTPIFDRTGVDEPQEFATFILNKILSSSDPNPKSLKLPRFTRDIGNLVRANFEISRQNITKLRRKIVSQLNGEEKDTHLSTFVVSYAYIIVSIVKAKQLEKKGKVGFLFSGDCRSRLNPPLPTNYFGNCVIGSFVLIDTETIAEGDLIVWVVKELSRMIKELEIRVKNMSSDREIENLLNFERNIVKVADLIIGVAGSPQFGIYGTDFGWGKPNKIQIVSIDNRGAISMAESRDGNGGIEVGLVLKKHEMEMFSSLFVNGFKDM
>Casp06589.t1
MKKNMVVSLISKETIKPHSSEPSKPFKLSLIDQITPTTYVPLIFFYPPKSNQNPQFSANLKLSLSQTLSTFFPFSGRLKDNLFVHNFDQGIPFSETQVSSELSEFLQNPQLEMLNQFLPTNPFTQITPPNETLPLIYVQFNVFRCGGIAIAICISHKIADTDSITKFLRNWADNNKNLGTGSSIITPSFEESSLRFLPFKSNPSHQISIMEKLFFNERKRTTRRFVFDAKAIISLKEKAKTEIGNGEKRTPTRVKTLSSFIWKCCTLASASISTTPRLSISAHFVNIRQRTQPCLPEYSIGNLLWCAVAVYDIVDSSDYTGTSDDLKHFVGLTEESVEEVNVEFLKSLEGEKGVEVVCEFLEQVNEIYSTNPEMFLFTSWLNFGINDLDFGLGKPAWVSLMGEVGPGLRNLTIFKELYDGIEAWITLDDDVMEILEHDPEFLEFATPNPPIFNYVKT
>Casp07007.t1
MASTPTPVKVTEVYQVTPPESASELTLSLTFFDTLWFKFHPVERIFFYQLTADSSFFYSSIIPQLKHTLSLTLLHFRLLAGNLIWPSDSPKPVILYTSNDAVSFTVAESDADFHLLSGATIHEAADSHHYIPELSVSETKAAIVSFQITLFPNKGFSIGVSSHHAISDGKSIIMFLKAWAHISKHNTSLLPPELTPFLDRTIIQDPEGIDMVYLNDWITLAAKMGTDSNPTSLKLIPITPPSDIKLVRGTFRLSGEDVKKLREKVISAIGEGNYSTFVLTYAYVAINMVKSKGIERNMKLGIGFMADCRSRLNPPVLDNYFGNCVIPHGLQTEAGLVMEEDGFVFLVGKLKETMKKIEKGVMDGAKDILSDYLKAEPDLMETIGVAGGNRFGVYGTDFGWGRPEMVEVTSVDRTGAFSMAETKHGNGVEIGLAMAQNQMEIFSSLFINGLK
>Casp07010.t1
MASPITVEILEICKVSPATDSNESGQRVSLPLTFLDTRLLKFPQAERLYLYKHINLTPTLFMSTLLPRLKRALSLTLLHFLPLAGHLTWPSDSPEPVILYDPNDAVSLTVAESNAEFDLLVSNGIREVAELRPYIPLLSMSDTEVSIISLRITLFPSKGFSVGIAMHHAALDGKSGSMFLKAWAYTCKQNQNDGKPSPLLPELIPSFDRSCIGDPDGLNSYYLNYWLTTTGSNSNSKTKTCSLKLLQNVTDFEFPPNLVRATFQLSPDYISKIRESVQNYYDFQNQPVKLHMSTFVIVSAYASVCLVKARGGSGGGGRMVYFLVAVDCRSRLDPPVPENYMGNCVFTVDMVAEAADFMEENGVAVIAKRISDLVKGLEKGVFEGARERHVRLRNAGEQVQKIAVVGSPRFCYYEEDFGWGKPEKVEITSVDRINGIQLKDSRDGNGVIEIGLIAYLLAQSTGSEAVTSPVWGLVWKLQVPERLCFFVWLVLHNGVLSNENCVRRQLCLDSSCSECEESDSILHIIHDCRATRAVWALWIHLEERNEFFSRPLAAWIQKNCW
>Casp08439.t1
MLKHSLSKTLSHFYPLAGKIKDSVSIECDDEGAYYTQATANISMPEFLRQPDHAALKKLVPDASLFNQNPIGAFILMIQETVFACGGLAIGFSVSHLANDGYGFGSFIKAWASAASSPTKQIPSPYLIDMFPLYEPFPKEALFPALWAPFAKKRKVAVRRFVFDASAIDKLKSEAASRVENPSRVEVVSAIISKSVKAALNAKSIAICHSANMRRKATPPIPDCSMGNFVWILQAICTEDESQVSEFVCKLRGAKRKLEREFMKNVEGGFSKLFEEVKEMGLALSSDDGMEMIGFSSLCNFDGYDGSDFGWGKPVWMSHCTAFDSEEIMFHNMGTLNDTRTGINGVEAWMYLYPEVASAMEKDVHLLQYASIDPPPLTMLHHHYS
>Casp10079.t1
MLMILQDPQVPQIEKLLSFNSQEMSSNEEILAAQVNHFDCGGIAISICIYHQIADGSTMASFATTWAAVASGKAATNIKGMIYDCTSIFPPQKTEGCSWNRFVEKDVLSNSMMKRFVFDRSKLEALREKVVNGSGLAYTGRPSRVETINAVIWGALLEITNLNTAIIAVNLRKRLIPPLSELSIGNIYHLVNCSMNENQIDYPDLVGKIHESIDIVKDDYVRKIHEGGRYFEFMKDVAEKPNLMMNVFAFSSWCRFPFYKVDFGWGNPIWVAPAVKVHNAAVVIDTKDGEGIEAWVGLPKEDMAKFQRIPHICSYASFNPTI
>Casp10082.t1
MAGRIKNNSYIDCNDEGAFYTEAHATGDMSMLLQDPQLPQIEKLLPFDSQELSSDEEILGVQVNHFDCGGMAISICIWHKIADGSSAASFATTWAAVASGEATTNIEGIIYDCTSIFPPQKTQGFLWNILPKKDALRNSMMKRFVFDHSDLVALREKVGNGSKLECLGRPSRVETVTAVIWGAVIESTDVNTTLIAVNLRKRLIPPLPELSVGNIYHTANCLENENQIDYNYLAGKIHESIGIVKDDYVRKIYEEGSYFEFMKDIVEKPNFMKNVFGFSSWCRFPFYEVDFGWGKPIWVTTTLKVNQGAILIDTKDGKGIEAWVTLPKENMAKFQQIPYICSYASFNPTIY
>Casp10418.t1
MPYFNCGGLAVSFAMPHMFGDALSTVMFLNCWAKIAQGKTNTNMIVDTHPIGSSSIFPPRDSSDSDIMQLNPKEKFVTKSFVFDAFTLSALKDKYCCCGEKLSRVQALSVFLLSRIAATIYQAKEGCNKNQSLVISAMNLRPLFDPPISKQSFGNFVWMGVAMIDNMEDMAKKDEYCCEISVRIKDSIKSVNKETVKKLQSDDQVNVLKETFAEAAKGEIDKYAFSSLCNFPMYEVDFGWGKPEWIVTTAMILDNVSTVFMDTKEGKGIEAWISMSEGDMAKSEIDNKLLSYLSSTPNGNQNIIKQSRL
>Casp10420.t1
MKIEVEFISEELIKPSSPTPHHLRRINNYSYTDCNDKGALFIEAKANCHLSDILQNRNQYHNHRFKFIPLQPHQGIHQYGSLFQITYFNCFGLALSFAMPHMLGDALSMVMFVNCWATIAQGKMDTNMFVDTHPIGSSSIFLPRDSSEAYLKQLPLKEKIVTKSFVFDAFTLSALKDKYSRHGENISRVQALSVFLLSRIAATISQAKEGCNMNRSVVVNAINLRPLVDPPISKQSFGNFLWMGVAMFDNMEDMAKEDEYCCEIAARIRDSIKKVAKGEIDKYAFSSLCNFPIYETDFGWGKPEWIVTTGLMLDNVYIIFIDTKEGKGIEAWISMSEEDMAKFESDNKLLSHLSSTLNGNQNLMKQSRL
>Casp10428.t1
MKIEVEFISKEFIKPSSPTPHHLHKLHFSFLDQIQAQISPPMTMFYEKSNIFTNLERCNLLKKSLSKALTIFYPLAGRINNYSYADCNDEGALFIEAKANCQLSDILQNRNQFHNHRFKFIPLQPHQGVHQYGSLFQITYFNCSGFALCFAMPHMLGDALSQVMFVNCWAAIALGNTNTDTFVDTHPIGSSSIFPPRDSSDSDLMQLVLKQKIVTKSFVLDAFTLSAFKDKYSRDGEKLSRVQALSVFLLSRIAAATKEDGNKNRSVVVNAMNLRPLVDPSISKQSFGNFVWMGVAMVDNMEDMAKEDEYCCEIGVRIKDSIKSANKETAKKLQSHDQLNFLKETFAEVAKGEIDKYAFSSLCNFPLYEVDFGWGKPEWITTTAMILDNVYIIFMDTKEGKGIEAWISMSEEDMAKFEIDNKLLSYLSSTSNGNQNLIKQSRL
>Casp14312.t1
MQEAIALHPYVPELPVSDSKAAIIAFQITFFPSQGFSVGISSHHAMFDGKSVAMFMKSWAHICKENFDIRALLPLELTPTFDRTVVEELQGLDMFYMKNIFSSSDSNPRSLKPPRVIRDMENLVRANYELSRQDITNLRRKIVSQQNQKEGEDSKRIHLSTFVLSYAYIIVTIVKAKNMDRKGKVGFLFAAECRSRLNPPLPMNYFGNCVVGSFGLVDTETVLDKNGTASVANELSQVIKGLETRVKNMSVEREIEQTLKLKNIMQVADVVIGVAGSPRFEIYGTDFGWGKPKKIQVVSIDRNVSISMAECRDSNGGLEVGLALKKSEMEMFTFLFANGLKYM
>Casp14488.t1
MEMFNMFLPYEPFCQQQLDPTLGMLAVQVNIFDCRGIALGICFSHKINDGITSSSFMKTWAANSCGSGSYNKVVQPMFPLQKSLPSRYTSLMETLWFSKKAFRTRRFVFNESAVSALRAKGRSEYVENTTRVEALSAFIWKCLMKACENISGSPRSSVMSQAVNIRRLMKPRMSRYSIGNLVWSTIARYNPDETEMEMKELVSLVREGVGKVNSEYLKSFSGEKGSLAILEHLDELGEISMENPDVFSFYSWHTFGFSEIDFGYGKPVWVGIFGEANRNSASDSNFIILKDVGRNNAIEAWMTLDENIMDVLENDPEFLAFASLNPCITSD
>Casp15327.t1
MPSAPDTLSLISKCTVLPHHKSSMDDLKLSVSDLPMLSCHYIQKGALFSRPPISIDSLLSILKRSLSQTLSQFPPLAGRLRTDSDGYIYITCNDAGVDFFHASATHLSVRDIISPLHVPDSVKSFFAFDRTVSYRGHFMPILAVQVTELADGVFIGCSCNHSVTDGTSFWNFFNTFAELSRGVNKISKQPDFTRNSVLISPAVLPLPAGGPKVTFNENEPLSERIFSFSREAILKIKAKANNKKWIENSGIDSAVELMGKQSNDTYLNHNNGNMMTAILETLFKTAAKPQEMETNGVSAVAIQAAAEISSFQSLCALLWRAVTRARKLNPTKTTTFRMAVNCRTRLNPKLDPLYFGNAIQSIPTYASAGDVTSRDLRWCAEQLNKNVVAHNDTTVRRFVNDWEANPRCFPLGNFDGASMTMGSSPRFPMYDNDFGWGRPVAVRSGRGNKFDGKISAFPGREGGGSVDLEVVLAPETMAGIESDSEFMQFVCN
>Casp16041.t1
MGKIQVDILSKEIIKPSSQTKIHHLKPFKFSLFNQLTPTTYTPIILFYPNINDPNITFSKIIPRLKLSLSQTLTFFYPFSGRVVDNLHIDHFNQGVPLIVAKVSGVGRLSDYLKNPEIEFLNRFLPSQPYFKEIDLGNIPQVAFQVNVFPCGGVVLGWVASHKLIDALSGSCFINAWASISKTGDLPEHMQPNCTQGSVFFPTKNPFPEEHLSLMESLWFTRDNYITRRFVFDAKTISSLRAKATGERDGKTIKPSRIEALTCFIWKCSMAASRAISGTPKASILVEALNLRTRTKPPMSSTSIGDIFWWATAIADASLENKELYELSFLLNDAINLYDIDYMKTLQGEEGFETMGDYCSQLHGLFSVEKPDIFAFTSWCHLGITKVDFGWGEPYWVGILGKAGPEFRNLTVFVDARDGNGIEAWITLDEQRMKYFECDPEFLAYACPNPKISCM
>Casp16815.t1
MASSDSHAAVRVTELCKITPPDSPTDSSLPLTFFDIFWLKFHPVERIFFYQLTSDSDSDSDSDSFSSDILPRLKQSLSLTLLHFIPFAGNLIWPSHSDKPFLLYSPGDGVSLTVAESDADFHTLAGDSIREIPDSHNYVPELNVTETEAATMSFQITLFPKKGYCIGISSHHGIFDGKSIIMFIKAWAHICKNGSSSLPPELTPVIDRAVIQDHQGLESLYLNSWMSLASLPDMDSNPRSLKLFNMPRSTSDTNYVRGTFKLSGEDIKKIRQKTFSQLDKEATNQANSKYFSTFVLSYAYVSITITKAKALEPNKHMILGFTADVRARQDPPISTNYFGNCVSINFTQHKVGIFIKEDGLAFVAEELTKLVKGLEKKGGLDGAKEKFEESMKMDPATTEGIGVAGSPRFEVYGIDFGFGKPIKVEITSIDRNTSISMAETRDGNGGVEIGVVLKKDEMEIFYSLFLYGLKDL
>Casp16817.t1
MALSDSPARIFFYQLTTDSDSDSSSFYSDILPKLKQSLSLTLLHFIPFAGNLIWPSHSDKPFLLYSPGDGVSLTVAESDADFHSLSGDSIREIPDSHHYIPELNVSETRASTMSFQVTVFPNQGFCIGISSHHAIFDGKSIIMFLKAWAHICKNDNGNPALPPELTPVHDRTLIQDPEGLEMLYLNNWMAVASLPGIETNTKSLKLFTMPRSTTDRSFVRGTFKLSDEHINKIRQKALSQLHEKATNQAISKHLSTFVLSYAYVSIAITKAKALEPNKKLVLGFTADVRARLDPHVPANYFGNCVSSNFIQTQVESLIKQDRLAFVVEELTKIIKGLENKGGLDGAKERFKAYMKMDPATIQRIGVARSPRFEVYGIDFGWGKHKKVEVTSIDKSTSISMAERRDGNGGVEIGVVLKKDEMKIFYSLFLDGLKDI
>Casp18114.t1
MKLKKTLEKLLEPYDFLAGRLKMNLETGRFEIDCNGAGAVFVSGFSECRLEELGDLVYPNPAFGQLIINNSDKEDSPLCIIQVTSFSCGGFAIGISTNHATFDGISFKKFLENIAAVAADKPLAVTPCNDRQLLAARSPPRITFPHPELLKLNTPPSHQQPDPAAAAVFDLTKENLDFKIFRLTSADISDLKEKAKSHPGARVSGFNVVTAHIWRCKALSYDENEDLERESRILYAVNIRPRLKPPLPDSYTGNAVLTAYASAKCKELKEGPVSRLVEMIVAGAKRMDDEYARSAIDWGEMNKGFPNGEFLVSSWWKLGFEQVVYPWGSPAFSCPLVYHRKDIILLFPDTVDREGVNVLVALPPQQMVKFQLLFKNFLLD
>Casp18441.t1
MVSTEIIKPLSSSKTHPTSHNFSFFDQICARIYVPYIFFYPIEELHHKYSSIQLKNSLSAALSLYYPLAGRFKDDLTIDCNDEGIEFQEARLDSDLSEILKNPNDEVIKILFPENLQYKDPISSSFLIVKVNLFNCGGIGLAICISHKVMDMAAFSYFINHWAGIARNSGDKFRPDFSSIPWLYPSIDLPVMECYEPDKLKCVSRRVVFHGSKIDELKLMVEKEVPNPTRFEVVTAVLYKSALAASTEFSNVVKSAVMHVGTNLREKVIPQFPESSSGNFSGTFSVSAGMEMEMETLTREIKKEKIKYWKCCGEKSDGEDLCRYVLEASVGLRRGHGKNEEGYLCSSYCRYPFYEADFGWGKPAWVSLASCEVKNVMIMLDTRDGDGIEAYVSLQEQVMEIFENDHFLLSFASINPSVLPEST
>Casp21349.t1
MPPVYFSTVFFYPNNNDFIVSQRSSVLKDSLSKTLSNFYPLAGKIKDYVSIDCNDEGAYYTEATANISMSEYLTQPDPKFIPKLVPDVSSLSQNPIGAFIVLIQVTTFACGGLVIGTSVSHMILDGISFSSFVKAWALAASGKQITPPDFDSRSLFPCYDGNIPKQALLPALWIPFLRKHKFAMRRVIFDASGIHQLKAKAAAEVENPSRVEVVSSIITKTVKAALNAKSLVITHPVNLRRKATPRFPDSSLGNFVWPTQAIVTEKEKQLVYNLRKAIRKVDSEFVKGAIEEGSSKLFQDMKEMGNAVPDNQDEMDLLAFGSWCNFGLYNNSDFGWGNPIWTAPSNDISDSEEITFNNMVCLNDTRSGDGVEAWLYLPQHVIAVVEKDDELLQYASIDLSPLNI
>Casp21353.t1
MNDGPSPPSFSGGFSGAMDIKEPRVCCEGRIAAWRKIKDYVSIDCNDEGAYYTEATANISMTEYLTQPDPKFIPKLVPDVSSLSQNPIGAFVVLIQVTTFACGGLVIGLSVSHMVLDGISFSSSIKAWALAASGKQITPPDFDSRSLFPCYDGNIPKQALFPALWIPFLQKHKFAMRRLIFDASGIDKLKAKAAAEVENPSRVEVVSSIVAKTVKSALNTKSLVINHAVNMRRKATPPFPDSSLGNFVRPTQAIITEKETQLVYNLRNAIRKVDSEFVKNAIEDGSSKLFQDMKEMGNAVSDNQDKIELLVFGSWCNFGLYHSSDFGWGNPIWISPSNDINDSEEITFNNMVCLVDTRSEDGVEAWLYLPQHVIAVVEKDGELLQYASIDLSPLNI
>Casp21549.t1
MPHMLGDALSMFIFLNGWAAVARGNNTVEEIPPFVADSIFPARSISGFDLTQWVFKENVVTKSFVFDAVTISSLKDKYSANGEKLSLVRALSVFLLSRIIAATNSQAKARCMVINSVNVRQILDPPISKNSFGNLIWAAAALIDDMDDIGTKEFEFVRRIKDSIKSVNAESLKKLQKGELNFTKKEYMESLKGEIVKYLFAGLSNFPVYEVDFGWGQPQWITTSGMIFDNLIVCLDTKDQKGIEAWISLTEEDMAKFENDQQLLSHLSSNPNFISRL
>Casp23354.t1
MIQETTFACGGIALGLSVSHMVLDATALASFIKAWAEDKEIPSSPNLESPSVFPQHDSFPKEAMFSVLWSPFFKKGKVAARRLMFKGSDIDKLKANSASSSVEKPSRVEVVSALISKSVMSAYNAPISIIHTINMRRKATPAFPESSVGNFIWIVGAALSEKERHLSSLVYKIRSAIRKVDSEFVKKIGALKLFREVKVIETELSKAQEYMSFTSWCNLGLYENSDFGWGKPIWIAQCSIIDSEDITFNNYCASCGER
>Casp28328.t1
MPCSSSSSPTIISKCTIYPDQNSDTKSLKLSVSDIPMLSCQYIQKGVLLSYPPYSFDDLITFLQNSLSSALSFFPALAGRLSTDSDGHIHILCNDAGIDFVQAKAKHISLLNILSPDLDVPYCFKEFFAFDKLLSYNGHFKPLVAVQVTELSDAVFIGCTVNHSVTDGTSFWHFFNTVAEICKGDTSTNKKISNLPDFSRNTVFNSPAVLKFPSGGPKVTFSGDEPLRERIFHFSRDAILKMKYKANNGNLLERKGKQNNDSWKIVNGESNGGVLKNKTDEISSFQSLCAQLWRCVTRARKLPLTKTTTFRMAVNCRHRLEPRLHPYYFGNAIQSIPTIAPIGELLSRDLHWGADLLHKNVVAHDDNTVRKGIADWEREPRLFPLGNFDGASITMGSSPRFPMYNNDFGWGRPLAIRSGRANKFDGKISAFPGKEGNGSVDLEVVLSPETMARIEQDEEFMQYVS
>Casp31115.t1
MPPIFYSTALFYPNSISQTSLVLKQSLSKTLTQFYLLAGKIKDYLSIDCNDEGALYIEAKSNIPLSSYLTQPDLTLLPKLVPDLSPFLQNPIGAFLLMIQQTTFTCGALALGLSVSHMALDGTAFASFIKAWALTASSAAKQLTNPNLDAPITFPQRNSFPKEALFSALWSPYLKKGKVIATRLMFVASALDKLKADAASSSVEKPSRVEVVSAVICKSVMAAFNAPLSITHAVNVRGKATPPFPNCSLGNFVWLVPVTFRETDGEVSSLVWKIRNAVRKVDSDFVKRIGGLEVFEEVKVIESEVSEGKEVMLVTSWCNFGVYKNSDFGWGNPTWIAPCSRIDSEEIEFNNACILNDTKIGNGIEAWVYLPQDVAAIVEKDAHLLQYASINPTPL
>Casp33290.t1
MEVEIISKQTLKPSSSSTNSHPKIFKNSLLDQLIPSRYASLVLFYPNIKFQNPDVFTRLNLLKTSLSETLACLYPLAGKIRDDLSIDCNDEGACFVETRVNCSLDEFLTEPDLLSLKKFLPCEIISDESVIGTFVTNVQVNVFKCDGIAVGICLSHRIMDGDGLRMFLKMWSGNSTNKSSSEMSNFAVSGAATLFPADDLLWLRDSSNAMWGSFLKKGNFVTKRFVFYASAIANLKDKAIESGIISRPTRVEVVSAILWKSMAAASAELNGFRRASLLNHLVNLRRRMEPSLVENTLGNFLWIASAKQDEAAEPEFGDLAGKIREAIARIDNEFVEKMKGKEGKCVMLKSLEEGGEIGGENGVDYFVFSSWCNFGYYDVGDFGWGEPVWVSGVGGVGGANSMFFNVIMLVDTKCGGGVEAWVTLDEQEMAILERDDDIIRFASLDLSPLLNL

Gene tree

Gene tree

Sheng jun & Dong yang Teams, Yunnan agriculture University